Subfamily Aleocharinae Fleming, 1821

 

Tribe Aleocharini Fleming, 1821

Subtribe Aleocharina Fleming, 1821

 

Genus Aleochara Gravenhorst, 1802

Subgenus Aleochara Gravenhorst, 1802

 

Aleochara (Aleochara) claviger Sharp

Aleochara claviger Sharp, 1874a: 7.

Aleochara niponensis Sharp, 1888: 281.

Aleochara (Aleochara) clavigera: Smetana, 2004: 353 [misspelling].

Aleochara (Baryodma) niponensis: Smetana, 2004: 354.

Korean records. Aleochara (Aleochara) claviger: Park and Ahn, 2009: 521 (South); Luo and Zhou, 2012: 191.

Distribution. Korea, China (Chongqing, Sichuan), Japan.

 

Aleochara (Aleochara) coreana Bernhauer

Aleochara coreana Bernhauer, 1926: 271.

Aleochara (Aleochara) lewisia Bernhauer, 1936: 324.

Korean records. Aleochara coreana: original description (North). Aleochara (Aleochara) coreana: Smetana, 2004: 353 (North); Park et al., 2011: 75 (North, South).

Distribution. Korea, China (Beijing, Heilongjiang), Japan.

 

Aleochara (Aleochara) curtula (Goeze)

Staphylinus curtula Goeze, 1777: 730.

Staphylinus brachypterus Geoffroy, 1785: 167.

Staphylinus limbatus Fabricius, 1801: 600.

Staphylinus fuscipes Gravenhorst, 1802: 93.

Aleochara puncticeps Thomson, 1860: 248.

Aleochara discoidea Sharp, 1874a: 7.

Korean records. Aleochara (Aleochara) curtula: Yuh et al., 1985: 252 (South); Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 30 (South); Paśnik, 2001: 230 (North); Cho et al., 2002: 36 (South); Cho and Kim, 2003: 126 (South); Smetana, 2004: 353 (North, South), Luo and Zhou, 2012: 193.

Distribution. Korea, China (Beijing, Chongqing, Gansu, Liaoning, Northeast Territory, Shaanxi, Sichuan), Japan, Russia (East Siberia, West Siberia), Morocco, Europe, Nearctic, Neotropical region.

 

Aleochara (Aleochara) kochi

BernhauerAleochara (Polychara) kochi Bernhaer, 1941: 211.

Aleochara (Aleochara) kochi: Smetana, 2004: 354.

Korean record. Aleochara (Aleochara) kochi: Park and Ahn, 2010b: 313 (South).

Distribution. Korea, Japan.

 

Aleochara (Aleochara) lata Gravenhorst

Aleochara lata Gravenhorst, 1802: 186.

Aleochara lata chinensis Bernhauer, 1933b: 48.

Korean records. Aleochara lata: Haku, 1936: 125 (South). Aleochara (Aleochara) lata: Yuh et al., 1985: 252 (South); Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 30 (South); Paśnik, 2001: 231 (North); Cho and Kim, 2003: 126 (South); Smetana, 2004: 354 (North, South).

Distribution. Korea, China (Jilin), Japan, Russia (East Siberia, West Siberia), Cyprus, Turkey, Europe, Afrotropical, Nearctic, Neotropical region.

 

Aleochara (Aleochara) parens Sharp

Aleochara parens Sharp, 1874a: 6.

Korean records. Aleochara (Aleochara) parens: Yuh et al., 1985: 252 (South); Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 31 (South); Paśnik, 2001: 231 (North); Cho and Kim, 2003: 126 (South); Smetana, 2004: 354 (North, South), Luo and Zhou, 2012: 198. Aleochara parens: Ishii, 1940: 43 (South); Kim and Nam, 1982: 150 (South).

Distribution. Korea, China (Beijing, Jiangsu, Liaoning), Japan.

 

Subgenus Coprochara Mulsant and Rey, 1874a

 

Aleochara (Coprochara) bipustulata (Linnaeus)

Staphylinus bipustulata Linnaeus, 1760: 232.

Staphylinus bipunctata Olivier, 1795: [42]: 31.

Aleochara nitida Gravenhorst, 1802: 97.

Aleochara cursor Stephens, 1832: 159.

Aleochara velox Stephens, 1832: 159.

Aleochara dorsalis Stephens, 1832: 160.

Homalota biguttula Kolenati, 1846: 8.

Baryodma nitida fusconotata Mulsant and Rey, 1874a: 439.

Baryodma nitida laetipennis Mulsant and Rey, 1874a: 439.

Baryodma nitida transita Mulsant and Rey, 1874a: 439.

Aleochara unicolor Schilsky, 1908: 600.

Korean records. Aleochara bipustulata: Kim, 1980: 199 (South). Aleochara (Coprochara) bipustulata: Yuh et al., 1985: 252 (South); Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 30 (South); Smetana, 2004: 355.

Distribution. Korea, China (Beijing), Russia (East Siberia, Far East, West Siberia), North Africa, Holarctic region.

 

Aleochara (Coprochara) squalithorax Sharp

Aleochara squalithorax Sharp, 1888: 282.

Korean records. Aleochara (Coprochara) squalithorax: Ahn et al., 2000: 244 (South); Cho and Ahn, 2001: 31 (South); Smetana, 2004: 356 (South); Frank and Ahn, 2011: 19 (South). Aleochara squalithorax: Park and Ahn, 2004: 196 (South).

Distribution. Korea, Japan.

 

Aleochara (Coprochara) verna Say

Aleochara verna Say, 1833: 58.

Aleochara languida Sachse, 1852: 117.

Baryodma (Coprochara) pauxilla Mulsant and Rey, 1874a: 443.

Aleochara alticola Sharp, 1883b: 148.

Aleochara (Coprochara) cedari Likovský, 1984: 8 [replacement name].

Baryodma minuta Casey, 1906: 161 [homonym].

Baryodma pumilio Casey, 1911: 6.

Baryodma tolerata Casey, 1911: 6.

Aleochara tecumseh Muona, 1977: 16 [replacement name].

Aleochara (Coprochara) tanumi Likovský, 1984: 8 [replacement name].

Korean records. Aleochara (Coprochara) verna: Paśnik, 2001: 231 (North); Smetana, 2004: 356 (North).

Distribution. Korea, China (Heilongjiang, Shaanxi, Xinjiang), Japan, Russia (Far East), North Africa, Palearctic, Nearctic region.

 

Subgenus Emplenota Casey, 1884a

 

Aleochara (Emplenota) fucicola Sharp

Aleochara fucicola Sharp, 1874a: 9.

Homalota variolosa Weise, 1877a: 89.

Korean records. Emplenota fucicola: Paśnik, 2001: 231 (North). Aleochara fucicola: Park and Ahn 2004: 195 (South). Aleochara (Emplenota) fucicola: Cho and Ahn, 2001: 30 (South); Smetana, 2004: 356 (North, South); Frank and Ahn, 2011: 19 (South); Yamamoto and Maruyama, 2012: 11 (North, South).

Distribution. Korea, China (Anhui, Heilongjiang, Hongkong), Japan, France.

 

Aleochara (Emplenota) puetzi (Assing)

Emplenota puetzi Assing, 1995: 225.

Korean records. Emplenota puetzi: Paśnik, 2001: 232 (North). Aleochara (Emplenota) puetzi:Smetana, 2004: 356 (North); Yamamoto and Maruyama, 2012: 18 (North).

Distribution. Korea, Japan, Russia (Far East).

 

Aleochara (Emplenota) segregata Yamamoto and Maruyama

Aleochara (Emplenota) segregata Yamamoto and Maruyama, 2012: 20.

Korean records. Aleochara (Emplenota) segregata: original description (South). Aleochara (Emplenota) fucicola: Ahn et al., 2000: 243 (South) [misidentification]. Aleochara puetzi: Park and Ahn, 2004: 196 (South) [misidentification]. Aleochara (Emplenota) puetzi: Frank and Ahn, 2011: 19 (South).

Distribution. Korea, Japan.

 

Subgenus Triochara Bernhauer, 1901b

 

Aleochara (Triochara) trisulcata Weise

Aleochara trisulcata Weise, 1877a: 88.

Korean records. Aleochara (Triochara) trisulcata: Ahn et al., 2000: 246 (South); Cho and Ahn, 2001: 31 (South); Smetana, 2004: 358 (South); Frank and Ahn, 2011: 20 (South); Yamamoto and Maruyama, 2012: 31 (South). Aleochara trisulcata: Park and Ahn, 2004: 196 (South).

Distribution. Korea, China (Hongkong), Japan.

 

Aleochara (Triochara) zerchei (Assing)

Triochara zerchei Assing, 1995: 231.

Korean records. Triochara zerchei: Paśnik, 2001: 232 (North).

Aleochara zerchei: Park and Ahn, 2004: 196 (South).

Aleochara (Triochara) zerchei: Smetana, 2004: 358 (North); Frank and Ahn, 2011: 20 (South); Yamamoto and Maruyama, 2012: 35 (South, North). Aleochara (Triochara) trisulcata: Ahn et al., 2000: 245 [misidentification].

Distribution. Korea, Japan, Russia (Far East).

 

Subgenus Xenochara Mulsant and Rey, 1874a

 

Aleochara (Xenochara) asiatica Kraatz

Aleochara asiatica Kraatz, 1859: 15.

Aleochara (Euryodma) asiatica: Bernhauer and Scheerpeltz, 1926: 780; Smetana, 2004: 356.

Aleochara (Polychara) asiatica: Cameron, 1939b: 644.

Aleochara (Xenochara) asiatica: Pace, 2001: 35.

Aleochara japonica Sharp, 1874a: 8.

Korean records. Aleochara (Xenochara) asiatica: Park and Ahn, 2010a: 23 (South).

Distribution. Korea, China (Hongkong), Japan, India (Himachal Pradesh, Uttaranchal, Uttar Pradesh), Nepal, Taiwan.

 

Aleochara (Xenochara) intricata Mannerheim

Aleochara intricata Mannerheim, 1830: 66.

Aleochara celeris Stephens, 1832: 161.

Aleochara terminata Stephens, 1832: 158.

Aleochara biguttata Heer, 1839: 315.

Aleochara (Baryodma) bipunctata var. croatica Penecke, 1901: 12.

Korean records. Aleochara (Baryodma) intricata: Paśnik, 2001: 230 (North); Smetana, 2004: 354 (North). Aleochara (Xenochara) intricata: Park and Ahn, 2010a: 25 (South).

Distribution. Korea, China (Heilongjiang), Russia (East Siberia, Far East, West Siberia), Palearctic region.

 

Aleochara (Xenochara) peninsulae Bernhauer

Aleochara (Polychara) peninsulae Bernhauer, 1936: 325.

Aleochara (Xenochara) peninsulae: Smetana, 2004: 360.

Korean record. Aleochara (Xenochara) peninsulae: Park and Ahn, 2010a: 27 (South).

Distribution. Korea, Japan.

 

Aleochara (Xenochara) puberula Klug

Aleochara puberula Klug, 1832: 139.

Aleochara vaga Erichson, 1839a: 172.

Aleochara deserta Erichson, 1839a: 173.

Aleochara decorata Aubé, 1850: 311.

Aleochara armitagei Wollaston, 1854: 559.

Aleochara badia Motschulsky, 1858c: 237.

Oxypoda sanguinolenta Motschulsky, 1858c: 241.

Oxypoda brunescens Motschulsky, 1858c: 243.

Aleochara dubia Fauvel, 1863: 428.

Oxypoda analis MacLeay, 1871: 135.

Baryodma bipartita Casey, 1893: 287.

Aleochara major Eichelbaum, 1912: 176.

Korean records. Aleochara (Aleochara) puberula: Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 31 (South). Aleochara (Xenochara) puberula: Smetana, 2004: 360 (South).

Distribution. Korea, China (Guangdong, Guangxi, Hongkong, Jilin, Liaoning, Yunnan), Japan, Cosmopolitan.

 

Aleochara (Xenochara) tristis Gravenhorst

Aleochara tristis Gravenhorst, 1806: 170.

Staphylinus bipunctata Olivier, 1795: 31.

Staphylinus geometrica Schrank, 1798: 642.

Aleochara bimaculata Stephens, 1832: 158.

Aleochara flavomaculata Ménétriés, 1832: 147.

Aleochara nigripes Miller, 1853: 27.

Aleochara erectesetosa Jekel, 1873: 41.

Baryodma (Polychara) discipennis Mulsant and Rey, 1874a: 360.

Korean records. Aleochara (Xenochara) tristis: Paśnik, 2001: 231 (North); Smetana, 2004: 361 (North).

Distribution. Korea, China (Heilongjiang), Russia (Caucasus, East Siberia, Far East, West Siberia), Europe, Northern Africa, Afrotropical, Nearctic, Oriental region.

 

Aleocharini incertae sedis

 

Genus Amarochara Thomson, 1858

Subgenus Amarochara Thomson, 1858

 

Amarochara (Amarochara) umbrosa (Erichson)

Calodera umbrosa Erichson, 1837: 304.

Homalota teuis Heer, 1839: 340.

Oxypoda funicularis Hochhuth, 1872: 104.

Nasirema parviceps Casey, 1893: 309.

Ischnopoda (turfanensis) turfanensis Pace, 1998b: 404.

Korean records. Amarochara umbrosa: Paśnik, 2001: 227 (North); Assing, 2010: 1140 (North). Amarochara (Amarochara) umbrosa: Smetana, 2004: 469 (North).

Distribution. Korea, China (Hebei, Xinjiang), Russia (West Siberia), Palearctic region.

 

Tribe Athetini Casey, 1910

Subtribe Athetina Casey, 1910

 

Genus Acrotona Thomson, 1859

 

Acrotona aterrima (Gravenhorst)

Aleochara aterrima Gravenhorst, 1802: 83.

Bolitochara fuscula Mannerheim, 1830: 84.

Aleochara mannerheimii C.R. Sahlberg, 1831: 380.

Aleochara brunnipes Stephens, 1832: 133.

Aleochara luripes Stephens, 1832: 136.

Aleochara obscurata Stephens, 1832: 137.

Homalota lugens Kiesenwetter, 1844: 319.

Atheta aterrima: Brundin, 1952: 121.

Korean records. Acrotona aterrima: Paśnik, 2001: 202 (North); Smetana, 2004: 362 (North).

Distribution. Korea, Russia (East Siberia, West Siberia), Kazakhstan, Uzbekistan, Europe.

 

Acrotona grata (Cameron)

Atheta (Acrotona) grata Cameron, 1933a: 216.

Acrotona perbella Brundin, 1952: 143.

Atheta (Xenota) grata: Yosii and Sawada, 1976: 24.

Korean records. Acrotona grata: Paśnik, 2001: 203 (North); Smetana, 2004: 363 (North).

Distribution. Korea, Japan, Russian (Far East), Taiwan.

 

Acrotona lutulenta (Sharp)

Homalota lutulenta Sharp, 1888: 293.

Korean records. Acrotona lutulenta: Paśnik, 2001: 203 (North); Smetana, 2004: 364 (North).

Distribution. Korea, Japan.

 

Acrotona paeksongricus Paśnik

Acrotona paeksongricus Paśnik, 2001: 205.

Korean record. Acrotona paeksongricus: original description (North).

Distribution. Korea. Acrotona setaria (Brundin)Atheta setaria Brundin, 1952: 124.

Korean records. Acrotona setaria: Paśnik, 2001: 204 (North); Smetana, 2004: 365 (North).

Distribution. Korea, Mongolia, Turkmenia.

 

Acrotona suspiciosa suspiciosa (Motschulsky)

Homalota suspiciosa Motschulsky, 1860b: 90.

Atheta (Coprothassa) anepsia Cameron, 1920a: 95.

Korean records. Acrotona suspiciosa: Paśnik, 2001: 204 (North).

Acrotona suspiciosa suspiciosa: Smetana, 2004: 365 (North).

Distribution. Korea, China (Beijing, Gansu, Guizhou, Shaanxi, Yunnan, Xinjiang), India (Uttaranchal, Uttar Pradesh), Sri Lanka, Oriental region.

 

Acrotona suyangsani Paśnik

Acrotona suyangsani Paśnik, 2001: 204.

Korean records. Acrotona suyangsani: original description (North); Smetana, 2004: 365 (North).

Distribution. Korea. Acrotona vicaria (Kraatz)

 

Homalota vicaria Kraatz, 1859: 38.

Homalota inornata Kraatz, 1859: 39.

Homalota peregrina Kraatz, 1859: 39.

Homalota termitophila Motschulsky, 1860b: 91.

Atheta (Acrotona) pseudoparens Cameron, 1933a: 215; Sawada, 1977: 198.Atheta (Acrotona) taedia Cameron, 1933a: 215; Sawada, 1977: 198.Atheta (Acrotona) cariei Pace, 1984: 263.Atheta (Acrotona) vicaria immixta Pace, 1986: 175.

Korean records. Acrotona vicaria: Paśnik, 2001: 204 (North). Acrotona inornata: Smetana, 2004: 364 (North).

Distribution. Korea, China (Beijing, Hongkong, Sichuan, Yunnan, Zhejiang), Japan, India (Darjeeling District, Sikkim, Uttaranchal, Uttar Pradesh), Nepal, Afrotropical, Oriental region.

 

Acrotona vivida (Sharp)

Homalota vivida Sharp, 1874a: 15.

Korean records. Acrotona vivida: Paśnik, 2001: 204 (North); Smetana, 2004: 366 (North).

Distribution. Korea, Japan.

 

Genus Adota Casey, 1910

 

Adota magnipennis (Bernhauer)

Atheta (Anopleta) magnipennis Bernhauer, 1943: 184; G. Benick, 1970: 94.

Adota magnipennis: Smetana, 2004: 366.

Korean records. Adota magnipennis: Park et al., 2007: 208 (South); Frank and Ahn, 2011: 20 (South).

Distribution. Korea, Japan.

 

Genus Amidobia Thomson, 1858

 

Amidobia talpa (Heer)

Homalota talpa Heer, 1841: 594.

Homalota parallela Mannerheim, 1844: 178.

Atheta (Microdota) talpa Brundin, 1948: 44.

Korean records. Amidobia talpa: Paśnik, 2001: 198 (North); Smetana, 2004: 370 (North).

Distribution. Korea, Russia (East Siberia, West Siberia), Europe.

 

Genus Amischa Thomson, 1858

Subgenus Amischa Thomson, 1858

 

Amischa (Amischa) bifoveolata (Mannerheim)

Bolitochara bifoveolata Mannerheim, 1830: 79.

Homalota cavifrons Sharp, 1869: 177.

Amischa vogti G. Benick, 1967: 24.

Korean records. Amischa cavifrons: Paśnik, 2001: 198 (North). Amischa bifoveolata: Smetana, 2004: 370 (North).

Distribution. Korea, Russia (East Siberia, Far East, West Siberia), Europe, Nearctic region.

 

Genus Atheta Thomson, 1858

Subgenus Atheta Thomson, 1858

 

Atheta (Atheta) basicornis (Mulsant and Rey)

Homalota basicornis Mulsant and Rey, 1852: 42.

Korean records. Atheta (Atheta) basicornis: Paśnik, 2001: 224 (North); Smetana, 2004: 373 (North).

Distribution. Korea, Russia (West Siberia), Scandinavia, Central Europe.

 

Atheta (Atheta) castanoptera (Mannerheim)

Bolitochara castanoptera Mannerheim, 1830: 77.

Aleochara xanthoptera Stephens, 1832: 116.

Aleochara consobrina Stephens, 1832: 122.

Aleochara foveatocollis Stephens, 1832: 134.

Homalota pertyi Heer, 1839: 329.

Homalota merdaria Thomson, 1852: 141.

Korean records. Atheta (Atheta) castanoptera: Paśnik, 2001: 224 (North); Smetana, 2004: 373 (North).

Distribution. Korea, Russia (Caucasus, East Siberia, West Siberia), Iran, Syria, Europe, North Africa.

 

Atheta (Atheta) euryptera (Stephens)

Aleochara euryptera Stephens, 1832: 135.

Homalota validicornis Märkel, 1845: 212.

Homalota succicola Thomson, 1852: 141.

Homalota cribrosa Mulsant and Rey, 1873b: 543.

Atheta (Atheta) euryptera japonica Bernhauer, 1907: 407.

Korean records. Ischnopoda (Atheta) euryptera japonica: Yuh et al., 1985: 251 (South). Atheta japonica: Kim et al., 1994: 144 (South). Atheta (Atheta) japonica: Cho and Ahn, 2001: 32 (South). Atheta (Atheta) euryptera: Paśnik, 2001: 224 (North); Smetana, 2004: 374.

Distribution. Korea, China (Zhejiang), Japan, Russia (East Siberia, Far East), Holarctic region.

 

Atheta (Atheta) graminicola (Gravenhorst)

Aleochara graminicola Gravenhorst, 1806: 176.

Aleochara moesta Zetterstedt, 1828: 101.

Aleochara coracina C.R. Sahlberg, 1830: 350.

Aleochara foveola Stephens, 1832: 124.

Stonalota granulata Mannerheim, 1846: 508.

Homalota nigrina Aubé, 1850: 304.

Homalota tenuicornis Thomson, 1852: 134.

Homalota flavicornis Gerhardt, 1891: 205.

Atheta (Metaxya) septentrionalis Poppius, 1909: 22.

Atheta graminicola stockleini G. Benick, 1938: 184.

Korean records. Atheta (Atheta) graminicola: Paśnik, 2001: 224 (North); Smetana, 2004: 374 (North).

Distribution. Korea, Russia (Caucasus, East Siberia, West Siberia), Kyrgyzstan, Mongolia, Europe, Nearctic region.

 

Atheta (Atheta) sauteri Bernhauer

Atheta (Atheta) sauteri Bernhauer, 1907: 407.

Atheta (Acrotona) reitteriana Bernhauer, 1939a: 109.

Atheta (Notothecta) reitteriana: Yosii and Sawada, 1976: 44.

Atheta (Atheta) cameroni Pace, 1987: 422.

Korean records. Notothecta reitteriana: Paśnik, 2001: 199 (North). Atheta (Atheta) sauteri: Smetana, 2004: 376 (North).

Distribution. Korea, China (Beijing, Guangdong, Guizhou, Hongkong, Jiangsu, Shaanxi, Sichuan, Zhejiang), Japan, India (Uttaranchal, Uttar Pradesh), Nepal, Taiwan.

 

Atheta (Atheta) taesongsanensis Paśnik

Atheta (Atheta) taesongsanensis Paśnik, 2001: 224.

Korean records. Atheta (Atheta) taesongsanensis: original description (North); Smetana, 2004: 376 (North).

Distribution. Korea.

 

Atheta (Atheta) transfuga (Sharp)

Homalota transfuga Sharp, 1874a: 13.

Atheta (Atheta) kubotai Bernhauer, 1943: 185.

Korean records. Atheta transfuga: Kim et al., 1994: 144 (South); Cho and Ahn, 2001: 32 (South).

Distribution. Korea, Japan.

 

Subgenus Badura Mulsant and Rey, 1873a

 

Atheta (Badura) nana (Kraatz)

Homalota nana Kraatz, 1859: 36.

Atheta (Datomicra) kanagawana Bernhauer, 1907: 399.

Atheta (Microdota) vulgaris Cameron, 1920c: 258.

Atheta (Acrotona) seticauda Bernhauer, 1935a: 12.

Atheta (Datomicra) nana: Cameron, 1939b: 387.

Atheta (Datomicra) pseudosordidula Cameron, 1944: 106.

Korean records. Atheta (Microdota) nana: Paśnik, 2001: 208 (North). Atheta (Badura) nana: Smetana, 2004: 377 (North).

Distribution. Korea, China (Guangxi, Hongkong, Sichuan), Japan, Taiwan, India (Uttaranchal, Uttar Pradesh).

 

Atheta (Badura) ririkoae Sawada

Atheta (Badura) ririkoae Sawada, 1989: 285; Smetana, 2004: 377.

Korean records. Atheta ririkoae: Park et al., 2007: 209 (South); Frank and Ahn, 2011: 21 (South).

Distribution. Korea, Japan.

 

Atheta (Badura) tokiokai (Sawada)

Ischnopoda (Dinaraea) tokiokai Sawada, 1971b: 306.

Atheta (Badura) tokiokai: Smetana, 2004: 377.

Korean records. Atheta tokiokai: Park et al., 2007: 210 (South); Frank and Ahn, 2011: 21 (South).

Distribution. Korea, Japan.

 

Subgenus Chaetida Mulsant and Rey, 1873b

 

Atheta (Chaetida) longicornis (Gravenhorst)

Aleochara longicornis Gravenhorst, 1802: 87.

Bolitochara validicornis Mannerheim, 1830: 83.

Aleochara antennata Stephens, 1832: 115.

Homalota socialis Lucas, 1849: 101.

Homalota tropica Motschulsky, 1858c: 256.

Korean records. Atheta (Chaetida) longicornis: Paśnik, 2001: 226 (North); Smetana, 2004: 378 (North).

Distribution. Korea, China (Sichuan), Russia (East Siberia, West Siberia), Palearctic, Oriental region.

 

Subgenus Coprothassa Thomson, 1859

 

Atheta (Coprothassa) melanaria (Mannerheim)

Oxypoda melanaria Mannerheim, 1830: 70.

Aleochara tenera C.R. Sahlberg, 1831: 386.

Homalota testudinea Erichson, 1839a: 131.

Homalota taeniata Kolenati, 1846: 7.

Phloeocharis gigantea Bernhauer, 1901a: 114.

Korean records. Atheta (Coprothassa) melanaria Paśnik, 2001: 207 (North); Smetana, 2004: 378 (North).

Distribution. Korea, China (Gansu, Shanxi, Yunnan), Japan, Russia (Caucasus, East Siberia, Far East, West Siberia), Altai, Mongolia, Europe, Oriental region.

 

Subgenus Datomicra Mulsant and Rey, 1873b

 

Atheta (Datomicra) celata (Erichson)

Homalota celata Erichson, 1837: 335.

Homalota montana Mulsant and Rey, 1852: 38.

Atheta arenicola Thomson, 1868: 321.

Homalota germana Sharp, 1869: 235.

Atheta (Datomicra) exsecta G. Benick, 1975: 14.

Korean records. Atheta (Datomicra) celata: Paśnik, 2001: 214 (North); Smetana, 2004: 379 (North).

Distribution. Korea, Russia (East Siberia, West Siberia), Algeria, Mongolia, Turkey, Uzbekistan, Europe.

 

Atheta (Datomicra) furtiva Cameron

Atheta (Dimetrota) furtiva Cameron, 1939b: 378.

Atheta (Datomicra) furtive fragilior Pace, 1986: 177.

Korean records. Atheta (Dimetrota) furtiva: Paśnik, 2001: 217 (North). Atheta (Datomicra) furtiva: Smetana, 2004: 379 (North).

Distribution. Korea, China (Sichuan, Yunnan), India (Darjeeling District, Himachal Pradesh, Sikkim), Nepal.

 

Atheta (Datomicra) lewisiana Cameron

Atheta (Datomicra) lewisiana Cameron, 1933a: 214.

Korean records. Atheta (Datomicra) lewisiana: Paśnik, 2001: 214 (North); Atheta (Datostiba) lewisiana: Smetana, 2004: 380 (North).

Distribution. Korea, China (Beijing, Hongkong, Jiangsu, Shaanxi, Shanxi, Yunnan, Zhejiang), Japan, Java, Nepal, Oriental region.

 

Atheta (Datomicra) nigra (Kraatz)

Homalota nigra Kraatz, 1856: 287.

Atheta (Datomicra) delecta G. Benick, 1975: 13.

Korean records. Atheta (Datomicra) nigra: Paśnik, 2001: 214 (North); Smetana, 2004: 379 (North).

Distribution. Korea, China (Yunnan), Canary Islands, Kazakhstan, Europe.

 

Atheta (Datomicra) pyongangsani Paśnik

Atheta (Datomicra) pyongangsani Paśnik, 2001: 215.

Korean records. Atheta (Datomicra) pyongangsani: original description (North); Smetana, 2004: 379 (North).

Distribution. Korea.

 

Atheta (Datomicra) sordidula (Erichson)

Homalota sordidula Erichson, 1837: 335.

Korean records. Atheta (Datomicra) sordidula: Paśnik, 2001: 214 (North); Smetana, 2004: 379 (North).

Distribution. Korea, China (Xinjiang), Russia (East Siberia, West Siberia), Palearctic, Oriental region.

 

Atheta (Datomicra) subsericans Cameron

Atheta (Atheta) subsericans Cameron, 1939b: 355.

Korean records. Atheta (Datomicra) subsericans: Paśnik, 2001: 215 (North); Smetana, 2004: 380 (North).

Distribution. Korea, China (Northeast Territory, Sichuan), India (Himachal Pradesh, Kashmir, Uttaranchal, Uttar Pradesh).

 

Subgenus Dimetrota Mulsant and Rey, 1873a

 

Atheta (Dimetrota) aeneipennis (Thomson)

Homalota aeneipennis Thomson, 1856: 103.

Homalota cursor Motschulsky, 1858c: 251 [homonym].

Homalota immunda Brisout de Barneville, 1860: 342.

Homalota cursoria Gemminger and Harold, 1868: 533 [replacement name].

Ischnopoda (Coproceramius) tenuiducta Sawada, 1970: 52.

Atheta (Dimetrota) laetipennis G. Benick, 1982: 98.

Korean record. Atheta (Dimetrota) aeneipennis: Smetana, 2004: 380 (North).

Distribution. Korea, Japan, Russia (East Siberia, West Siberia), Iran, Mongolia, Uzbekistan, Europe, Oriental region.

 

Atheta (Dimetrota) altaica Bernhauer

Atheta (Dimetrota) altaica Bernhauer, 1901a: 109.

Korean records. Atheta (Dimetrota) altaica: Paśnik, 2001: 216 (North); Smetana, 2004: 381 (North).

Distribution. Korea, Russia (West Siberia, North European Territory), Central Asia, Northern Scandinavia, Nearctic region.

 

Atheta (Dimetrota) atramentaria (Gyllenhal)

Aleochara atramentaria Gyllenhal, 1810: 408.

Bolitochara impressifrons Mannerheim, 1830: 84.

Aleochara borealis C.R. Sahlberg, 1831: 361.

Aleochara consimilis Stephens, 1832: 116.

Aleochara atricornis Stephens, 1832: 133.

Aleochara aenescens Zetterstedt, 1838: 77.

Homalota morio Heer, 1839: 336.

Homalota transfuga Sharp, 1874a: 13.

Atheta (Metaxya) geysiri Schubert, 1909: 774.

Atheta atramentaria var. spuria Eichelbaum, 1913: 148.

Atheta (Dimetrota) sulputrida Cameron, 1933a: 214.

Atheta bowringi Bernhauer, 1936: 317.

Atheta (Dimetrota) sublaevana Cameron, 1939b: 379.

Atheta (Notothecta) kunmingensis Pace, 1993: 106.

Korean records. Atheta atramentaria: Kim et al., 1994: 144 (South). Atheta (Atheta) atramentaria: Cho and Ahn, 2001: 31 (South). Atheta (Dimetrota) atramentaria: Paśnik, 2001: 216 (North); Smetana, 2004: 381 (North).

Distribution. Korea, China (Beijing, Guangxi, Hongkong, Hubei, Sichuan, Yunnan), Japan, Russia (East Siberia, Far East, West Siberia), Palearctic, Afrotropical, Oriental region.

 

Atheta (Dimetrota) chagangensis Paśnik

Atheta (Dimetrota) chagangensis Paśnik, 2001: 221.

Korean records. Atheta (Dimetrota) chagangensis: original description (North); Smetana, 2004: 381 (North).

Distribution. Korea.

 

Atheta (Dimetrota) machonryongica Paśnik

Atheta (Dimetrota) machonryongica Paśnik, 2001: 222.

Korean records. Atheta (Dimetrota) machonryongica: original description (North); Smetana, 2004: 382 (North).

Distribution. Korea.

 

Atheta (Dimetrota) myohyangsani Paśnik

Atheta (Dimetrota) myohyangsani Paśnik, 2001: 218.

Korean records. Atheta (Dimetrota) myohyangsani: original description (North); Smetana, 2004: 382 (North).

Distribution. Korea.

 

Atheta (Dimetrota) munsteri Bernhauer

Atheta (Dimetrota) munsteri Bernhauer, 1902b: 55.

Korean records. Atheta (Dimetrota) munsteri: Paśnik, 2001: 217 (North). Boreophilia munsteri: Smetana, 2004: 396 (North).

Distribution. Korea, Scandinavia, Nearctic region.

 

Atheta (Dimetrota) nigripes (Thomson)

Homalota nigripes Thomson, 1856: 102.

Homalota villosula Kraatz, 1856: 305.

Atheta (Dimetrota) nigripes: Brundin, 1953: 427; Benick and Lohse, 1974: 209.

Korean records. Atheta (Dimetrota) nigripes: Paśnik, 2001: 217 (North); Smetana, 2004: 382 (North).

Distribution. Korea, Europe.

 

Atheta (Dimetrota) photaechonica Paśnik

Atheta (Dimetrota) photaechonica Paśnik, 2001: 220.

Korean record.Atheta (Dimetrota) photaechonica: original description (North); Smetana, 2004: 383 (North).

Distribution. Korea. Atheta

 

Atheta (Dimetrota) weisei Bernhauer

Atheta (Atheta) weisei Bernhauer, 1907: 408.

Ischnopoda (Coproceramius) multispina Sawada 1970: 51.

Korean records. Atheta weisei: Cho et al., 2002: 35 (South); Atheta (Dimetrota) weisei: Paśnik, 2001: 218 (North); Smetana, 2004: 383 (North).

Distribution. Korea, China (Jilin), Japan.

 

Subgenus Microdota Mulsant and Rey, 1873b

 

Atheta (Microdota) amicula (Stephens)

Aleochara amicula Stephens, 1832: 132.

Aleochara picipennis Stephens, 1832: 132.

Homalota longula Heer, 1839: 334.

Homalota sericea Mulsant and Rey, 1852: 41.

Homalota subsericea Wollaston, 1864: 540.

Homalota jezabel Saulcy, 1865a: 438.

Microdota terricola Mulsant and Rey, 1873b: 351.

Homalota meludyi Quedenfeldt, 1884: 366.

Atheta (Microdota) amiculaattarum Bernhauer, 1929b: 201.

Atheta (Microdota) nuda G. Benick, 1975: 4.

Korean records. Atheta (Microdota) amicula: Paśnik, 2001: 207 (North); Smetana, 2004: 384 (North).

Distribution. Korea, China (Beijing), Russia (West Siberia), Palearctic, Neotropical region.

 

Atheta (Microdota) formicetorum Bernhauer

Atheta (Microdota) formicetorum Bernhauer, 1907: 400.

Korean records. Atheta (Microdota) formicetorum: Paśnik, 2001: 208 (North); Smetana, 2004: 385 (North).

Distribution. Korea, Japan.

 

Atheta (Microdota) hamgyongsani Paśnik

Atheta (Microdota) hamgyongsani Paśnik, 2001: 211.

Korean records. Atheta (Microdota) hamgyongsani: original description (North); Smetana, 2004: 385 (North).

Distribution. Korea.

 

Atheta (Microdota) kangsonica Paśnik

Atheta (Microdota) kangsonica Paśnik, 2001: 209.

Korean records. Atheta (Microdota) kangsonica: original description (North); Smetana, 2004: 386 (North).

Distribution. Korea.

 

Atheta (Microdota) kobensis Cameron

Atheta (Microdota) kobensis Cameron, 1933a: 212.

Korean records. Atheta (Microdota) kobensis: Paśnik, 2001: 208 (North); Smetana, 2004: 386 (North).

Distribution. Korea, Japan.

 

Atheta (Microdota) koreana Bernhauer

Atheta koreana Bernhauer, 1923: 128.

Atheta (Amidobia) bulbosa Sawada, 1974: 179.

Korean records. Atheta koreana: original description (North); Kim et al., 1994: 144. Ischnopoda (Atheta) koreana: Yuh et al., 1985: 251. Atheta (Microdota) koreana: Cho and Ahn, 2001: 32; Paśnik, 2001: 208 (North); Smetana, 2004: 386 (North).

Distribution. Korea, China (Liaoning), Japan.

 

Atheta (Microdota) palleola (Erichson)

Homalota palleola Erichson, 1837: 333.

Korean records. Atheta (Microdota) palleola: Paśnik 2001: 209 (North); Smetana, 2004: 387 (North).

Distribution. Korea, Russia (Caucasus, East Siberia), Europe.

 

Atheta (Microdota) scrobicollis (Kraatz)

Homalota scrobicollis Kraatz, 1859: 31.

Homalota inutilis Kraatz, 1859: 35.

Phloeopora indica Kraatz, 1859: 42.

Homalota cava Fauvel, 1875a: 738.

Microdota sericata Mulsant and Rey, 1875b: 182 [homonym].

Atheta sericina Fenyes, 1920: 186.

Korean records. Atheta (Microdota) scrobicollis: Paśnik, 2001: 209 (North); Smetana, 2004: 388 (North).

Distribution. Korea, China (Southeastern Territory), Japan, Ceylon, India (Darjeeling District, Himalchal Pradesh, Sikkim), Iran, Nepal, Singapore, Turkey, Europe, Afrotropical, Oriental region.

 

Atheta (Microdota) silvatica Bernhauer

Atheta (Microdota) silvatica Bernhauer, 1907: 405.

Korean records. Atheta (Microdota) silvatica: Paśnik, 2001: 209 (North); Smetana, 2004: 388 (North).

Distribution. Korea, Japan.

 

Atheta (Microdota) sogamensis Paśnik

Atheta (Microdota) sogamensis Paśnik, 2001: 210.

Korean records. Atheta (Microdota) sogamensis: original description (North); Smetana, 2004: 388 (North).

Distribution. Korea. 

 

Atheta (Microdota) subcrenulata Bernhauer

Atheta (Microdota) subcrenulata Bernhauer, 1907: 403.

Korean records. Atheta (Microdota) subcrenulata: Paśnik, 2001: 209 (North); Smetana, 2004: 388 (North).

Distribution. Korea, China (Beijing, Zhejiang), Japan.

 

Subgenus Oreostiba Ganglbauer, 1895

 

Atheta (Oreostiba) jangangsani Paśnik

Atheta (Oreostiba) jangangsani Paśnik, 2001: 213.

Korean records. Atheta (Oreostiba) jangangsani: original description (North); Smetana, 2004: 389 (North).

Distribution. Korea.

 

Atheta (Oreostiba) sparreschneideri Münster

Atheta sparreschneideri Münster, 1922: 207.

Atheta (Philhygra) anisi Bernhauer, 1913a: 134.

Korean records. Alaobia sparreschneideri: Paśnik, 2001: 199 (North). Atheta (Oreostiba) sparreschneideri: Smetana, 2004: 389 (North).

Distribution. Korea, Europe.

 

Subgenus Orientatheta Sawada, 1985

 

Atheta (Orientatheta) chinkiangensis Bernhauer

Atheta (Coprothassa) chinkiangensis Bernhauer, 1939b: 145.

Korean records. Notothecta chinkiangensis: Paśnik, 2001: 198 (North). Atheta (Orientatheta) chinkiangensis: Smetana, 2004: 390 (North).

Distribution. Korea, China (Jiangsu), Japan.

 

Subgenus Pachyatheta Münster, 1925

 

Atheta (Pachyatheta) mortuorum Thomson

Atheta mortuorum Thomson, 1867: 281.

Atheta pseudocribrata Hanssen, 1935a: 34.

Atheta (Datomicra) pseudocribrata Hanssen, 1935b: 54. [homonym]

Atheta (Microdota) mortuorum: Brundin, 1948: 42; Palm, 1970: 192.

Korean records. Atheta (Microdota) mortuorum: Paśnik, 2001: 208 (North). Atheta (Pachyatheta) mortuorum: Smetana, 2004: 390 (North).

Distribution. Korea, Russia (East Siberia, West Siberia), Palearctic region. 

 

Subgenus Rhagocneme Münster, 1922

 

Atheta (Rhagocneme) dwinensis Poppius

Atheta (Dimetrota) dwinensis Poppius, 1908: 17.

Atheta (Rhagocneme) fractipes Münster, 1922: 206.

Korean records. Atheta (Dimetrota) dwinensis: Paśnik, 2001: 2176 (North); Smetana, 2004: 393 (North).

Distribution. Korea, Russia (North European Territory), Finland, Norway, Sweden.

 

 

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