Chekcklist of the Coastal Staphylinidae (Coleoptera) 


J. H. Frank1 and Kee-­Jeong Ahn2


1Entomology and Nematology Department, University of Florida, Gainesvill, FL 32611-0630, USA;

2Department of Biology, Chungnam National University,Daejeon 305-764, South Korea




Because this work deals with coastal species, primary geographical entries are given according to the oceans and seas on which species are found. Secondary entries are the names of the countries they inhabit, and tertiary entries (if any) are the (mainly political) subunits of larger countries, or islands belonging to the former. Compression of this information into a checklist required the use of abbreviations, which are as follows:


Codes used for oceans have 3 letters: ACO (Arctic Ocean), INO (Indian Ocean), NAO (North Atlantic Ocean), NPO (North Pacific Ocean), SAO (South Atlantic Ocean), and SPO (South Pacific Ocean). Names of seas and gulfs are spelled out: Andaman Sea, Arabian Sea, Arafura Sea, Bali Sea, Baltic Sea, Bering Sea, Bismarck Sea, Black Sea, Caribbean Sea, Celebes Sea, East China Sea, East Sea [sometimes called Sea of Japan, but that name is disputed (Wikipedia 2010)], Gulf of California (sometimes called Sea of Cortez), Gulf of Mexico, Irish Sea, Java Sea, Mediterranean Sea (here including Adriatic, Aegean, Ionian, Ligurian and Tyrrhenian Seas), North Sea, Sea of Okhotsk, Philippine Sea, Red Sea, South China Sea, Sulu Sea, Tasman Sea, and Timor Sea. Names of seas are not used throughout. We have used the name of the ocean in the broad sense (of which the sea is part) in instances where the name of the sea is not apparent from the literature. For example, some species known from New Zealand may be known from the west coast (the Tasman Sea), but if that was not apparent from the literature, we ascribed them to SPO (the South Pacific Ocean).


Country codes have 2 letters and are the International Standards Organization (ISO) abbreviations. They are given in parentheses. Those used are: AG=Antigua and Barbuda, AL=Albania, AR=Argentina, AU=Australia, BB=Barbados, BE=Belgium, BG=Bulgaria, BM=Bermuda, BR=Brazil, BS=Bahamas, CA=Canada, CL=Chile, CN=China, CO=Columbia, CU=Cuba, CY=Cyprus, DE=Germany, DJ=Djibouti, DK=Denmark, DM=Dominica, DO=Dominican Republic, DZ=Algeria, EC=Ecuador, EE=Estonia, EG=Egypt, ER=Eritrea, ES=Spain, ET=Ethiopia, FI=Finland, FJ=Fiji, FP=French Polynesia, FR=France, GB=Great Britain, GD=Grenada, GE=Georgia, GH=Ghana, GL=Greenland, GP=Guadeloupe, GR=Greece, HR=Croatia, HT=Haiti, ID=Indonesia, IE=Ireland, IL=Israel, IN=India, IQ=Iraq, IS=Iceland, IT=Italy, JM=Jamaica, JP=Japan, KE=Kenya, KN=St. Kitts Nevis, KP=North Korea, KR=South Korea, KY=Cayman Island, LB=Lebanon, LC=St. Lucia, LK=Sri Lanka, LY=Libya, MA=Morocco, MG=Madagascar, MM=Myanmar, MR=Mauritania, MS=Montserrat, MT=Malta, MU=Mauritius, MX=Mexico, MY=Malaysia, NA=Namibia, NC=New Caledonia, NG=Nigeria, NL=Netherlands, NO=Norway, NZ=New Zealand, PE=Peru, PG=Papua New Guinea, PH=Philippines, PL=Poland, PR=Puerto Rico, PT=Portugal, RE=Reunion, RO=Romania, RU=Russian Federation, SA=Saudi Arabia, SC=Seychelles, SD=Sudan, SG=Singapore, SN=Senegal, SE=Sweden, SO=Somalia, TH=Thailand, TN=Tunisia, TR=Turkey, TT=Trinidad and Tobago, TW=Taiwan, TZ=Tanzania, UA=Ukraine, UK=United Kingdom, US=USA, UY=Uruguay, VE=Venezuela, VI=US Virgin Islands, VN=Vietnam, WS=Samoa (formerly Western Samoa, not American Samoa), YE=Yemen, YU=former Yugoslavia, ZA=South Africa.


Where places within countries are mentioned, they are given after a colon (:) following the abbreviation of the country name, and either are spelled out or are abbreviated. For the USA and Canada, the abbreviations are the 2-letter postal codes (BC=British Columbia, NB=New Brunswick, NL=Newfoundland and Labrador, NS=Nova Scotia, NT=Northwest Territories, PE=Prince Edward Island, QC=Quebec, YT=Yukon Territory); for Mexico they have 2 letters (BN=Baja California, BS=Baja California Sur, CA=Campeche, CH=Chiapas, GU=Guerrero, JA=Jalisco, MI=Michoacán, NA=Nayarit, OA=Oaxaca, QR=Quintana Roo, SI=Sinaloa, SO=Sonora, TB=Tabasco, TM=Tamaulipas, and VC=Veracruz); for Japan, designations are for major islands and island groups: (HK=Hokkaido, HN=Honshu, KY=Kyushu, RY=Ryukyu, SH=Shikoku). For Great Britain (GB) they are England, N. Ireland, Scotland, and Wales.


Checklist    Aleocharinae    Others    References A-E    References F-O    References P-Z


Acknowledgements  This work was partilly supported by a grant from the National Institute of Biological Resources (NIBR), funded by the Ministry of Environment (MOE) and by Basic Science Research Program through the National Research Foundation of Korea (NRF) funded by the Ministry of Education, Science and Technology.





Coastal Staphylinidae. A) Aleochara zerchei on a sandy beach in Donghae, Korea; B) Aleochara sp. on a sandy beach in Donghae, Korea; C) Bryothinusa koreana under a stone on rocky headland in Dangjin, Korea; D) Atheta tokiokai on a sandy beach in Jejudo Island, Korea; E) Paramblopusa borealis under a stone on pebble beach in Alaska, USA; F) Diaulota aokii with barnacles on a rocky shore in Baeksu, Korea; G) Diaulota aokii with fresh seaweeds on a rocky shore in Jejudo Island, Korea; H) Larva of Diaulota aokii with barnacles on a rocky shore in Baeksu, Korea. 

Coastal Staphylinidae. A) Phucobius simulator on a sandy beach in Guryongpo, Korea; B) Liusus hilleri on a sandy beach in Donghae, Korea; C) Overwintering staphylinine species (Cafius histrio, Liusus hilleri and Philonthus nudus) under a wooden board on a sandy beach in Jindo Island, Korea; D) Larva of Cafius sp. under fresh seaweeds on a sandy beach in Jejudo Island, Korea; E) Cafius bistriatus on a sandy beach in North Carolina, USA; F) Cafius seminites under decaying seaweeds on a sandy beach in California, USA; G) Cafius rufescens on a sandy beach in Jindo Island, Korea; H) Philonthus nudus under decaying seaweeds on a sandy beach in Jindo Island, Korea.